Fillmore makes organic fuel and is sort of a hippie. He plays what he calls the "classics" and lives in a brightly coloured tent. He often at night goes out to look at the traffic light that only blinks yellow. He is very friendly and says that the "gas companies that feed us lies" to help promote his business. He also is a Volkswagen. In Cars 2, he is part of Lightning McQueen's pit crew.
- His System shape is different.
- His DUPLO version doesn't have any mirrors.